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Research
Interests
Although
I am interested in the whole field of modern evolutionary biology,
my principal research has been focused on two broad areas: evolutionary
ecology and insect systematics. My evolutionary ecology studies
have been concerned with speciation and adaptation of island arthropods
to exploit harsh or inhospitable environments, and the colonization
and impacts of alien species on island ecosystems. Both studies
have provided insights into the evolution of community structure
and ecosystem functioning. Additionally, studying the adaptations
of organisms to such inhospitable environments leads to a better
understanding of evolution (especially natural selection) in general.
These studies have resulted in the discovery of three new ecosystems
in Hawaii: the lava tube, lava flow aeolian, and montane aeolian.
The methods used include (1) experimental ecological studies,
wherein a controlled modification of a single parameter is made
in the field and the results compared with similar control sites;
(2) collaboration with colleagues on comparative physiological,
behavioral, ecological, and morphological studies between cave-adapted
organisms and their close relatives living in above-ground habitats;
and (3) detailed bioclimatic studies correlated with adaptations
to the environment by cave species. I enjoy the opportunity to
collaborate on the transferal of these methods to other ecological
and evolutionary studies that are of interest to colleagues and
students.
I
believe such studies should be based on a firm understanding of
systematics and have cultivated a strong foundation in biosystematic
and collection-based research in entomology. The proper care and
preservation of voucher specimens of published research of both
field and laboratory studies in evolutionary biology is of paramount
importance in the interpretation and lasting value of such research.
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Selected
Publications
| Hoch,
H. & F.G. Howarth. 1999. Multiple cave invasions
by species of the planthopper genus Oliarus in
Hawaii (Homoptera: Fulgoroidea: Cixiidae). Zoological
Journal of the Linnean Society. 127(4): 453-475. |
| Howarth,
F.G. 2000. Non-target effects of biological control
agents. Pp. 369-403. IN: G.M. Gurr & S.D.
Wratten, eds. Measures of Success in Biological
Control. Kluwer Academic Pup., Dordrecht. 448
pp. |
| Bax,
N., Carlton, J.T., Mathews-Amos, A., Haedrich,
R.L, Howarth, F.G., Purcell, J.E, Rieser, A. and
Gray, A. 2001. The Control of Biological Invasions
in the World's Oceans. Conservation Biology 15:
1234-1246. |
| Taiti,
S. & F.G. Howarth. 1998. Terrestrial isopods (Crustacea,
Oniscidea) from Hawaiian caves. Memoires de Biospeologie
24:97-118. |
| Hoch,
H. & F.G.Howarth. 1993. Evolutionary Dynamics
of behaviorial divergence among populations of
the Hawaiian cave-dwelling planthopper Oliarus
polyphemus (Homoptera: Cixiidae). Pacific Science
47:303-318. |
| Howarth,
F.G. 1993. High-stress subterranean habitats and
evolutionary change in cave-inhabiting arthropods.
Amer. Naturalist 142 (Suppl.):65-77. |
| Howarth,
F.G. & W.P. Mull. (1992). Hawaiian Insects and
their Kin. Times Editions, Singapore and UH Press,
Honolulu. 160 pp. |
| Howarth,
F.G. & G.W. Ramsay. 1991. The conservation of
island insects and their habitats. pp. 71-107.
IN Collins, N.M. & J.A. Thomas (eds.). The Conservation
of Insects and their Habitats. 15th Symposium
of the Royal Entomological Society of London.
Academic Press. |
| Howarth,
F.G. 1991. Environmental impacts of classical
biological control. Annual Review Entomology 36:
485-509. |
| Howarth,
F.G. and F.D. Stone. 1990. Elevated carbon dioxide
levels in Bayliss Cave, Australia: Implications
for the evolution of obligate cave species. Pacific
Science. 44: 207-218. |
| Howarth,
F.G. 1991. Environmental impacts of classical
biological control. Annual Review Entomology 36:
485-509. |
| Howarth,
F.G. 1987. Evolutionary ecology of aeolian and
subterranean habitats in Hawaii. Trends in Ecol.
Evol. 2: 220-223. |
| Howarth,
F.G. 1985. The impacts of alien land arthropods
and mollusks on native plants and animals. pp.
149-179. IN C.P. Stone & J.M. Scott (eds.) Hawai'i's
Terrestrial Ecosystems: Protection and Management.
University of Hawaii Press, Honolulu. 584p. |
| Howarth,
F.G. 1983. Bioclimatic and geologic factors governing
the evolution and distribution of Hawaiian cave
insects. Entomol. Gen. 8:17-26. |
| Howarth,
F.G. 1983. Ecology of cave arthropods. Annual
Review Entomol. 28: 365-89. |
| Ahearn,
G.A. & F.G. Howarth. 1982. Physiology of cave
arthropods in Hawaii. J. Exp. Zool. 222:227-38. |
| Howarth,
F.G. 1980. The zoogeography of specialized cave
animals: A bioclimatic model. Evolution 34:394-406. |
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